BULLETIN DE L’INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE

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BRUXELLES 2003 BRUSSEL

yWTUM Sey,

BULLETIN DE L’INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE

BULLETIN VAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN

SCIENCES DE LA TERRE AARDWETENSCHAPPEN VOL. 73

BRUXELLES 2003 BRUSSEL

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Rédacteur en chef - Hoofdredacteur - Editor. Annie V . D h o n d t

Secrétaire de rédaction - Redactiesecretaris - Associate editor: Jacques G o d e f r o id

Comité de rédaction - Redactiecomité - Editorial board : Pierre B u l t y n c k

Daniel C a h e n Michel D e l ie n s

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D e n is e B r ic e (L il le , F ra n c e )Jenaro L. G a r c i a - A l c a l d e (Oviedo, Spain)

M ic h a e l A . K a m in s k i (L o n d o n , U K )Jordi M a r t in e l l (Barcelona, Spain)

David B. W e i s h a m p e l (Baltimore, USA)

La rédaction remercie pour lecture critique de manuscrits:De redactie dankt volgende reviewers voor medewerking aan dit volume:

For this volume the following reviewers are gratefully acknowledged:

F. Alvarez, W. Bambach, A. Balinski, H. Blumenstengel, Denise Brice, R.G. Bromley, P. Copper, J. Day, S.K.S. Donovan, J. Fedorowski, H. Jahnke, A. Kroh, B. Mamet, Margaret Mamet, M. Mancenido,

B. Mistiaen, R. Mooi, W.A.Oliver, E. F. Owen, L. Popov, G. Rabeder, G. Racki,S. Rodriguez, G. Stringer, A. Turner.

BULLETINDE L’INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE

SCIENCES DE LA TERRE

BULLETINVAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURW ETENSCHAPPEN

AARDW ETENSCHAPPEN

Vol. 73 - 2003 ISSN 0374-6291

Publié, verschenen .published: 31.111.2003

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TABLE DES MATIÈRES CONTENTS

C a s ie r , J.-G. & P r é a t , A. - O stracods and rock facies associated w ith the D evonian - C arboniferous boundary series in the Aves- nois, N orthern France 83

C o e n - A u b e r t , M. - D escription o f a few rugose corals from the G ivetian Terres d ’Haurs Form ation in Belgium 11

G o d e f r o id , J. & H a u s e r , J. - The Frasnian Pentam erida and A trypida (B rachiopoda) from the R eichle quarry (Eifel, G erm any) 53

Jagt, J. W. M. - N ote on a M aastrichtian cidarid echinoid from Eijsden (Lim burg,The N etherlands) 149

J a g t , J. W. M. - The ichnofossil genera Ra- dulichnus and Renichnus in the M aastrichtian o f The N etherlands and B elgium 175

M o t t e q u in , B. - Two new atrypid brachio- pod species from the late Frasnian o f B el­gium

N o l f , D. - Fish otoliths from the Santonian o f the Pyrenean faunal province, and an over-

69

view o f all o to lith-docum ented N orth A tlantic L ate C retaceous teleosts 155

P o t y , E. & H e c k e r , M. R. - Parallel ecolution in European rugose corals o f the genus Lon,s- daleia M cCoy, 1849 (Low er Carboniferous) 109

R a d w a n s k a , U. & R a d w a n s k i , A. - Bos­qu et’s (1862) inarticulate brachiopods : D is­cinisca elslooensis sp. n. from the Elsloo C onglom erate 185

S a n t i , G., R o s s i , M. & P o m o d o r o , S. - Cave bear rem ains from the Buco d e ll’Orso Cave (Lom bardy - Italy). Part III - M orphom etrie analysis o f m etapodial bones 195

S a r m ie n t o , G. & B u l t y n c k , P. - Late O rdo­vician C onodonts from the Fosses Form ation, C ondroz Area, Belgium 5

S a r t e n a e r , P. - Tullypothyridina, new late G ivetian rhynchonellid (brachiopod) genus 29

S im o n , E. - A new Late M aastrichtian rhyn- chonellide brachiopod from the Kunrade L i­m estone facies near M aastricht (Southern Lim burg, The N etherlands) 137

BULLETIN DE LTNSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE BULLETIN VAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN

SCIENCES DE LA TERRE, 73: 5-9. 2003 AARDW ETENSCHAPPEN, 73: 5-9, 2003

Late Ordovician Conodonts from the Fosses Formation, Condroz Area, Belgium

by G raciela SA R M IEN TO & Pierre BULTYN CK

S a r m i e n t o , G. & B u l t y n c k , P., 2003. Late Ordovician conodonts from the Fosses Formation, Condroz area, Belgium. Bulletin de l'In ­stitut royal des Sciences naturelles de Belgique, Sciences de ¡a Terre, 73:5-9, I pi., 1 fig., Bruxelles - Brussel, March 3 1 ,2 0 0 3 .- ISSN 0374- 6291.

Abstract

Taxonomic study o f a conodont faunule obtained from the Bois des Presles Member o f the Fosses Formation, permitted identification o f Amorphognathus sp. cf. A. ordovicicus B r a n s o n & M e h l , Amorphog- nathus? sp., Panderodus gracilis ( B r a n s o n & M e h l ) , Birksfeldial sp., and Plectodina'l sp. This conodont association, attributed to the Amor­phognathus ordovicicus Biozone (Ashgill), is tentatively referred to the British Province o f the North Atlantic conodont Realm.

Key-words: Amorphognathus ordovicicus Biozone, conodonts, Ash­gill, North Atlantic Realm, Belgium.

Resumen

El estudio taxonómico de una faúnula de conodontos proveniente del Miembro Bois des Presles de la Formación Fosses ha permitido la identificación de Amorphognathus sp. cf. A. ordovicicus B r a n s o n & M e h l , Amorphognathus? sp., Panderodus gracilis ( B r a n s o n & M e h l ) , Birksfeldial sp. y Plectodina'l sp. Esta asociación de conodontos se atribuye a la Biozona de Amorphognathus ordovicicus (Ashgill) y se adscribe tentativamente a la Provincia Británica del Dominio Nord- Atlántico de conodontos.

Palabras Clave: Biozona de Amorphognathus ordovicicus, conodon­tos, Ashgill, Dominio Nord-Atlántico, Bélgica.

Résum é

L ’étude systématique d ’une faunule à conodontes, provenant du Mem­bre du Bois des Presles (Formation de Fosses) a permis l'identification des taxa suivants: Amorphognathus sp. cf. A. ordovicicus B r a n s o n & M e h l , Amorphognathus'! sp., Panderodus gracilis ( B r a n s o n & M e h l ) , Birksfeldial sp. et Plectodinal sp. Cette faunule à conodontes, appartenant à la Biozone à Amorphognathus ordovicicus (Ashgill), est provisoirement attribuée à la Province Britannique du Domaine à conodontes de 1’ Atlantique Nord.

Mots-clefs: Biozone à Amorphognathus ordovicicus, conodontes, Ash­gill, Domaine de l'Atlantique Nord, Belgique.

Introduction

In a preliminary sedimentological and palaeontological study of the carbonate beds from the Fosses Formation (Condroz Area, Belgium) T o u r n e u r et al. (1993, p. 675) mention the very rare presence of conodonts and figure a specimen of Panderodus sp. In the present paper a more significant conodont fauna from the same area is described. It was obtained from the Bois de Presles Member of the Fosses Formation in 1977 and 1983. Two discontinuously exposed sections were measured and sampled (Fig. 1).

In section 1 at Cocriamont two 5 kg samples were taken and only sample 1 at the base of the formation produced four conodont elements.

In section 2, south-east o f Presles and temporarily exposed along the access road to a new residential area, three 2 to 4 kg samples were taken from which sample 2 produced thirteen conodont elements.

The reader is referred to V e r n ie r s et al. (2002, pp. 24- 27) for the Ordovician lithostratigraphy used herein.

C onodont collection and biostratigraphic and palaeobiogeographic rem arks

Only few and not well preserved conodont elements were recovered. All the specimens exhibit a homogeneous Col­our Alteration Index (CAI) of 4.5, indicating a thermal interval of 190-320°C ( E ps t e in et a i, 1 9 7 7 ). Most of the elements are poorly preserved and fragmentary but defor­mations have not been observed. Due to this preservation state most of the specimens have been identified in open nomenclature. The conodont association is dominated by simple cones of Panderodus gracilis. The conodont collec­tion is housed in the Department of Paleontology o f the Royal Museum of Natural Sciences o f Belgium.

Although most of the conodonts from the Bois des Presles Member of the Fosses Formation have been iden­tified in open nomenclature, the conodont association is most likely indicative of the Amorphognathus ordovici­cus Biozone by the occurrence o f Amorphognathus sp. cf. A. ordovicicus and Birksfeldial sp.

Late Ordovician conodont provincialism was discussed

6 Graciela SARMIENTO & Pierre BULTYNCK

PRESLESBRUSSELS

LE ROUX Charleroi

50 km

km 2

Bois de Chaumont

km 26

COCRIAMONTSART- \

EUSTACHE i

500 m

Sections continue

Mbr Fm

green sandy shale with dark spots FLT

- 10 alternation of calcareous

_ g shale and shelly beds

Sample Presles 2

FOSL0 m calcareousshale BDP

thick bed of bioclastic limestone with rounded calcareous-phosphatic clasts ; unconformity at the base ;

marl-stone

0 m -L-ö-Sf-s-tst-Sample Cocriam ont 4-

Fig. 1 — Upper part: location o f the studied outcrops; the black rectangle on the inset map shows location o f the Presles Cocriamont area.Lower part: Schematic columnar sections o f localities 1 (Cocriamont) and 2 (Presles); FOS = Fosses Formation; BDP = Bois des Presles Member; FLT = Faulx-les-Tombes Member.

Late Ordovician Conodonts from the Fosses Formation, Condroz Area, Belgium 7

by S w e e t & B e r g s t r ö m (1984) and updated in terms of multielement taxonomy by B e r g s t r ö m (1990). Accord­ing to these authors, three different provinces, Baltic, British and Mediterranean, can be recognized for the North Atlantic Realm. The characteristic conodont taxa o f the Mediterranean Province have not been recorded in our collection, while the genera Birksfeldia and Plecto- dina that are common in the Baltic and British provinces occur in the Belgian association. Although the small size o f the studied conodont collection enable a conclusive assignment to one o f these two provinces, the presence of several elements o f Plectodinal sp., that is common in Great Britain but rare in Baltoscandia, suggests major affinities with the British Province.

T axon om ic rem arks

Genus Amorphognathus B r a n s o n & M e h l , 1933

Type species: Amorphognathus ordovicicus B r a n s o n & M e h l , 1933

Amorphognathus sp. cf. A. ordovicicus B r a n s o n & M e h l , 1933

PI. 1, Figs. 1-3

R e m a r k s

Fragmentary pectiniform (Pa and Pb) elements of the Amorphognathus apparatus are here compared with those o f A. ordovicicus. The distinction of A. ordovicicus from the older A. superbus (R h o d e s ) is mainly based on the M (holodontiform) element. In the same way, other Late Ordovician species of Amorphognathus, such as, A. lind­stroemi ( S e r p a g l i) and A. ventilatus F e r r e t t i & B a r n e s have been distinguished upon the base of the M element, that is not represented in our collection.

For the position of the elements within the apparatus we follow the notation proposed by A r m s t r o n g et a!. (1996).

The index species Amorphognathus ordovicicus has been recorded widely in Europe and North America.

Amorphognathus? sp.PI. 1, Fig. 8

R e m a r k s

One incomplete Pb? element with prominent basal ribs is assigned to this genus in open nomenclature.

Genus Panderodus E t h in g t o n , 1959

Type species: Paltodus unicostatus B r a n s o n & M e h l , 1933

Panderodus gracilis ( B r a n s o n & M e h l , 1933)PI. 1, Figs. 7, 9

R e m a r k s

Several almost complete specimens are present in our

collection. The elements are slender, sub-symmetrical, with an antero-lateral costa on each side. According to the reconstruction and the notation proposed by S a n s o m et al. (1995) for the genus Panderodus, they are identified as sub-symmetrical graciliform elements.

Panderodus gracilis is known from the Ashgill of Europe, North America and Asia. Its total stratigraphie range is from the Middle Ordovician into the Silurian.

Genus Birksfeldia O r c h a r d , 1980

Type species: Birksfeldia circumplicata O r c h a r d , 1980

Birksfeldial sp.PI. 1, Fig. 6

R e m a r k s

One incomplete Sc? element is tentatively attributed to this genus. According to O r c h a r d (1980, p. 19) the Sc element o f Birksfeldia has an anticusp and sharp anterior and posterior edges which are developed as broad keels. Also the presence of a short denticulate posterior process and a broad carina on the inner face o f the cusp was pointed out by this author. In our specimen the anticusp is shorter than that illustrated by O r c h a r d (1980, pi. 6, fig. 21), and the posterior process is only partially pre­served.

O r c h a r d ( 1980) erected this genus upon the base o f a Late Ordovician (Cautleyan-Rawtheyan) conodont col­lection from England and Wales. It has been also reported from the attenuated Dent Group (mid-to late Rawtheyan) in Cumbria (UK) by A r m s t r o n g et al. (1996). In the Holy Cross Mountains (Poland) D z ik (1999) identified the genus Birksfeldia within the interval o f the Amor­phognathus ordovicicus Biozone. F e r r e t t i (1998) illu­strated one Pb element of Birksfeldial sp. from the Prague Basin, (Bohemia). Scarce and not well preserved speci­mens that could correspond to this genus have been found in the Iberian Peninsula (S a r m ie n t o , unpublished).

Genus Plectodina S t a u f f e r , 1935

Type species: Prioniodus aculeatus S t a u f f e r , 1935

Plectodinal sp.PI. 1, Figs. 4-5

R e m a r k s

Fragmentary ramiform elements with compressed denti­cles and apparently well developed cusp are attributed to this genus in open nomenclature.

The distinction o f Plectodina S t a u f f e r from the most closely related genus Aphelognathus B r a n s o n et al., 1951 is based on some features that cannot be observed in our material. Based on the aspect of the basal cavity of the Sa element we considérer that these specimens could be part of the apparatus of Plectodina.

8 Graciela SARMIENTO & Pierre BULTYNCK

References

A r m s t r o n g , H., Jo h n s o n , E.W. & S co rr, R.W., 1996. Con­odont biostratigraphy of the attenuated Dent Group (Upper Ordovician) at Hartley Broughton in Furness, Cumbria, UK. Proceedings o f the Yorkshire Geological Society, 51: 9-21. B e r g s t r ö m , S.M., 1990. Relations between conodont provin­cialism and the changing palaeogeography during the Early Palaeozoic. Geological Society o f America Memoir, 12: 105- 121 .

B r a n s o n , E.B. & M ehl, M.G., 1933. Conodont studies. Uni­versity o f Missouri Studies, 8, Parts 1-4: 1-343.B r a n s o n , E.B., M e h l, M.G., & B r a n s o n , C.C., 1951. Rich­mond conodonts o f Kentucky and Indiana. Journal o f Paleon­tology', 25 : 1-17.Dzik, J., 1999. Evolution of the Late Ordovician high-latitude conodonts and dating o f Gondwana glaciations. Bolletino della Societá Paleontológica Italiana, 37: 237-253.E p ste in , A.G., E p ste in , J.B. & H a r r is , L.D., 1977. Conodont color alteration - An index to organic metamorphism. United States Geological Survey Professional Paper, 995: 1-20. E th in g to n , R.L., 1959. Conodonts o f the Ordovician Galena Formation. Journal o f Paleontology, 33: 257-292.F e r r e t t i , A., 1998. Late Ordovician conodonts from the Prague Basin, Bohemia. In: S z a n ia w sk i, H. (ed.), Proceedings o f the Sixth European Conodont Symposium (ECOS VI). Palaeonto- logia Polonica, 58: 123-139.O r c h a r d , M. J., 1980. Upper Ordovician conodonts from England and Wales. Geológica et Palaeontologica, 14: 9-44.

S an so m , I.J., A r m s t r o n g , H.A. & Sm ith , M.P., 1995. The apparatus architecture o f Panderodus and its implications for coniform conodont classification. Palaeontology, 37: 781-800.

S t a u f f e r , C.R., 1935. Conodonts o f the Glenwood Beds. Geo­logical Society o f America, Bulletin. 46: 125-168.

S w e e t, W.C. & B e r g s t r ö m , S.M., 1984. Conodont provinces and biofacies o f the Late Ordovician. Geological Society o f America, Special Papers, 196: 69-87.

V e r n ie r s , J., H e rb o s c h , A., V a n g u e s ta in e , M., G e u k e n s , F., D e lc a m b re , B., P in g o t, J.-L., B e la n g e r , I., H e n n e b e r t , M., D e b a c k e r , T., S in tu b in , M. & D e v o s, W„ 2002. Cambrian Ordovician - Silurian lithostratigraphic units (Belgium). Geo­lógica Belgica, 4 (1-2): 5-38.

Graciela S a rm ie n to Departamento de Paleontología,Facultad de Ciencias Geológicas, CSIC-UCM,E-28040 Madrid, Spain.E-mail: gsarmien@geol.ucm.es

Pierre B u l ty n c kKoninklijk Belgisch Instituut voor Natuurwetenschappen Vautierstraat 29, B-1000 Brussels, Belgium.E-mail: Pierre.Bultynck@naturalsciences.be Typescript submitted: September 15, 2002 Revised typescript received: November 20, 2002

Explanation of Plate 1

Specimens Figs. 1-3 are from sample Presles 2, specimens Figs. 4 to 9 from sample Cocriamont I .

Figs. 1-3 — Amorphognathus sp. cf. A. ordovicicus B r a n s o n & M e h l ; 1-Pa element, upper view, x 69; 2- Pa element, upper view, X 84; 3- Pb element, lateral view, x 137; IRScNB specimens N° b4l79, b 4 180, b 4 181.

Figs. 4-5 — Plectodinal sp.; 4- Sa element, posterior view, x 227; 5- S? element, posterior view, x 227; IRScNB specimens N°b4182, b 4 183.

Fig. 6 — Birksfeldial sp.; Sc? element, inner lateral view, x 167; IRScNB specimen N° b4184.

Figs. 7, 9 — Panderodus gracilis ( B r a n s o n & M e h l) ; 7a- graciliform element, lateral view, x 190; 7b- detail o f the microstructurenear the basal margin of the element, x 455; 9- graciliform element, lateral view, x 190; IRScNB specimens N° b4185, b 4 187.

Amorphognathus1 sp.; Pb? element, antero-lateral view, x 227; IRScNB specimen N° b4186.

Late Ordovician Conodonts from the Fosses Formation, Condroz Area, Belgium 9

P late 1