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ABSTRACT!HBC$CDEG:$;GG<=CC=G$DS@$;CA=IDC$@K$=I@J@LBI;J$DH=@<:$G=M=J@A=G$A<BF;<BJ:$B>

/@<DH$&F=<BI;$;>G$=Z;FB>=G$DH=C=$DH=@<B=C$ECB>L$;>$&ECD<;JB;>$A;CC==$;C$;$F@G=J

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ABSTRACT ................................................................................................................................................. II

DEDICATION ............................................................................................................................................ III

ACKNOWLEDGEMENTS ....................................................................................................................... III

CHAPTER 1: INTRODUCTION ............................................................................................................... 1

-#/#*&+$)'#*'(#6 ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U1!30.$14#,(#1 ?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U1#/1(!('(!.$!)$"&%(!&!$2*&-Q#/!&!()/???????????????????????????????????????????????????????????????????????????????? YQ&(/!#/&/,#$)2$,))4#*&!('#$%*##0(/- ??????????????????????????????????????????????????????????????????????????????? X+(!#*&!3*#$,(!#0????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U^

CHAPTER 2. THE DEMOGRAPHY OF BROWN TREECREEPERS IN CONNECTED ANDISOLATED, LARGE AND SMALL, WOODLAND PATCHES........................................................... 26

&%1!*&,! ????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Vg(/!*)03,!()/???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? VgQ#!")01??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Vh*#13+!1????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? YW0(1,311()/??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? YV&,R/)6+#0-#Q#/!1?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Y^+(!#*&!3*#$,(!#0????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Yh!&%+#1??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ]W2(-3*#$,&4!()/1 ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ]V

CHAPTER 3. EXPERIMENTAL EVIDENCE OF DISRUPTED DISPERSAL CAUSINGDECLINE OF AN AUSTRALIAN PASSERINE IN FRAGMENTED HABITAT.............................. 45

&%1!*&,! ????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ]^(/!*)03,!()/???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ]^Q#!")01??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ]c*#13+!1????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ^U0(1,311()/??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ^Y&,R/)6+#0-#Q#/!1?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ^c+(!#*&!3*#$,(!#0????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ^X!&%+#1??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? g]2(-3*#$,&4!()/1 ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? g^

CHAPTER 4. EFFECTS OF LANDSCAPE PATTERNS ON DISPERSAL SUCCESS: ASIMULATION OF BROWN TREECREEPER POPULATION DYNAMICS IN A SPATIALLYREALISTIC LANDSCAPE....................................................................................................................... 69

&%1!*&,! ????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? gX(/!*)03,!()/???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? hWQ#!")01??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? hV*#13+!1????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? hc0(1,311()/??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? cW&,R/)6+#0-#Q#/!1?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? cY+(!#*&!3*#$,(!#0????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? c^!&%+#1??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? cX2(-3*#$,&4!()/1 ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Xg

M

CHAPTER 5. HABITAT AND LANDSCAPE EFFECTS ON BROWN TREECREEPERDISTRIBUTION: INDEPENDENT EFFECTS OF WOODLAND LOSS AND FRAGMENTATIONAT MULTIPLE SCALES........................................................................................................................ 113

&%1!*&,! ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UUY(/!*)03,!()/?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UUYQ#!")01????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UUg*#13+!1??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UVW0(1,311()/????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UVU&,R/)6+#0-#Q#/!1???????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UV^+(!#*&!3*#$,(!#0??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UVg!&%+#1????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UYW2(-3*#$,&4!()/1 ????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UYg

CHAPTER 6. HABITAT VARIATION AND COOPERATIVE BREEDING OF THEAUSTRALIAN BROWN TREECREEPER .......................................................................................... 142

&%1!*&,! ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U]VQ#!")01????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U]^*#13+!1??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U]c0(1,311()/????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U]c&,R/)6+#0-#Q#/!1???????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U^Y+(!#*&!3*#$,(!#0??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U^^!&%+#1????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? U^c2(-3*#$,&4!()/1 ????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UgV

CHAPTER 7. CONCLUSIONS ............................................................................................................. 169

1./!"#1(1 ?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UgX23!3*#$*#1#&*,"??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? UgX&'(&/$Q&/&-#Q#/!$(/$!"#$/#6$#/0+&/0$!&%+#+&/01 ?????????????????????????????????????????????? UhU

CIRRICULUM VITAE............................................................................................................................ 173

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9:$J=CC$DH;>$VWW\F$L;AC?$$,@>>=ID=G$A;DIH$CB[=$S;C$F=;CE<=G$;C$DH=$D@D;J$;<=;$@K$DH=$C=D

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BC@J;D=G$A;DIH$H;G$;$I@>>=IDBMBD:$CB[=$=NE;J$D@$BDC$A;DIH$CB[=5$SH=<=;C$HBLHJ:$I@>>=ID=G

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V8?$$$($EC=G$DHBC$F=;CE<=$D@$GBMBG=$DH=$A;DIH=C$B>D@$DS@$I;D=L@<B=CT$I@>>=ID=G$7I@>>=ID=G

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7x$YWW$H;$;>G$w$YWW$H;8$SBDHB>$=;IH$A;DIH$I;D=L@<:$72BLE<=$V\V8?

Demographic data - 2<@F$UXXg\UXXc5$KB=JG$;CCBCD;>DC$;>G$($F@>BD@<=G

;AA<@ZBF;D=J:$^W$L<@EAC$B>$UX$J@I;DB@>C$B>$;$D@D;J$@K$Ug$A;DIH=C$DH<@ELH@ED$DH=$/=S

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@K$;GEJDC$;>G$c^_$@K$@KKCAL$S=<=$9;>G=G$SBDH$E>BNE=$I@J@<$I@F9B>;DB@>C$;>G$;$F=D;J

9;>G$K<@F$DH=$&ECD<;JB;>$%B<G$;>G$%;D$%;>GB>L$1IH=F=?$$#;IH$:=;<5$K<@F$J;D=$OEJ:$D@

=;<J:$0=I=F9=<5$S=$MBCBD=G$=;IH$D=<<BD@<:$;D$J=;CD$@>I=$=M=<:$U^$G;:C?$$&D$=;IH$MBCBD$S=

BG=>DBKB=G$=;IH$9B<G$A<=C=>D$D@$G=D=<FB>=$L<@EA$CB[=$;>G$K@JJ@S=G$K=F;J=C$D@$G=D=<FB>=

SH=DH=<$;$>=CD$S;C$A<=C=>D?$$-<@EA$F=F9=<C$9LB>L$>=CD$F;D=<B;J$D@$;$D<==$H@J=

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>=CDJB>LC?$$6H=>$>=CDC$S=<=$;II=CCB9J=$7VU_$@K$;JJ$>=CDC85$>=CD$CD;DEC$S;C$G=D=<FB>=G$9:

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KJ=GLJB>L?

Statistical analyses$\$($EC=G$;$2BCH=<`C$#Z;ID$D=CD$D@$G=D=<FB>=$SH=DH=<$DH=

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*#13+!1

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CHAPTER 4. EFFECTS OF LANDSCAPE PATTERNS ON

DISPERSAL SUCCESS: A SIMULATION OF BROWN

TREECREEPER POPULATION DYNAMICS IN A SPATIALLY

REALISTIC LANDSCAPE

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*#13+!1

Proportional sensitivity analysis - !H=$A=<K@<F;>I=$@K$DH=$F@G=J$S;C$<@9ECD$D@

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Mean population size – (>BDB;J$L<@EA$CB[=$;>G$L<@EA$A<@GEIDBMBD:$GBG$>@D$GBKK=<

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GBCA=<C=<$F@<D;JBD:$<;D=$7!;9J=$]\]5$F;D<BZ$=KK=ID8?

Movement between fragmented and contiguous habitat$r$($G=KB>=G$I@>DBLE@EC$;>G

K<;LF=>D=G$H;9BD;D$9;C=G$@>$I@>>=IDBMBD:5$H@S=M=<$;$CF;JJ$A=<I=>D;L=$@K$9B<GC$79=DS==>

Y\U]_8$CEII=CCKEJJ:$GBCA=<C=G$9=DS==>$I@>DBLE@EC$;>G$K<;LF=>D=G$H;9BD;D$GEL$=;IH

hX

<E>$72BLE<=$]\X8?$$6H=>$($<=CD<BID=G$KE<DH=<$;>;J:C=C$@K$GBCA=<C;J$=M=>DC$7B?=?5$K=F;J=

GBCA=<C;J$F@M=F=>DC$;>G$J@>L\GBCD;>I=$F@M=F=>DC8$D@$@>J:$DH@C=$GBCA=<CB>L$9B<GC$DH;D

GBG$>@D$I<@CC$9=DS==>$I@>DBLE@EC$;>G$K<;LF=>D=G$H;9BD;D5$<=CEJDC$GBG$>@D$GBKK=<?

!H=<=K@<=5$($<=A@<D$<=CEJDC$ECB>L$;JJ$CEII=CCKEJ$GBCA=<C;J$=M=>DC$B>$DH=$K@JJ@SB>L$;>;J:C=C?

Female dispersal distances - !H=$;M=<;L=$F;ZBFEF$GBCA=<C;J$GBCD;>I=$@K$K=F;J=C

GBG$>@D$GBKK=<$9=DS==>$K<;LF=>D=G$;>G$I@>DBLE@EC$H;9BD;D$72BLE<=$]\UW87!;9J=$]\]5

J@I;DB@>$=KK=ID85$:=D$S;C$L<=;D=<$SBDH$DH=$/=BLH9@<$F@M=F=>D$<EJ=$7!;9J=$]\]5$F@M=

=KK=ID8$;>G$SBDH$,@>CD;>D$F@<D;JBD:$<;D=C$7!;9J=$]\]5$F;D<BZ$=KK=ID8?$$)>J:$<E>C$SBDH$DH=

F@CD$C=M=<=$F;D<BZ$=KK=ID$71D=AVW8$A<@GEI=G$F;ZBFEF$GBCA=<C;J$GBCD;>I=C$;C$CH@<D$;C

DH@C=$@9C=<M=G$B>$DH=$KB=JG$72BLE<=$]\UW8?

Long-distance dispersal events – 1H@<D\GBCD;>I=$GBCA=<C;J$=M=>DC$S=<=$;JS;:C

F@<=$K<=NE=>D$DH;>$J@>L\GBCD;>I=$GBCA=<C;J$=M=>DC?$$!H=$A<@A@<DB@>$@K$J@>L\GBCD;>I=

F@M=F=>DC$S;C$HBLH=<$B>$K<;LF=>D=G$H;9BD;D$DH;>$B>$I@>DBLE@EC$H;9BD;D$7!;9J=$]\]5

J@I;DB@>$=KK=ID8$72BLE<=$]\UU8?$$!HBC$GBKK=<=>I=$S;C$F@CD$;AA;<=>D$SBDH$DH=$*;>G@F

F@M=F=>D$<EJ=$7!;9J=$]\]5$J@I;DB@>{F@M=$=KK=ID8$;>G$DH=$,@>CD;>D$GBCA=<C;J$F@<D;JBD:

<;D=$7!;9J=$]\]5$J@I;DB@>{F;D<BZ$=KK=ID85$;>G$J=;CD$;AA;<=>D$SBDH$;>$=ZD<=F=$F;D<BZ$=KK=ID

71D=A$VW8$72BLE<=$]\UU8?

Proportion of solitary males that recruit a female – 1@JBD;<:$F;J=C$B>$I@>DBLE@EC

H;9BD;D$S=<=$;JS;:C$F@<=$JBP=J:$D@$<=I<EBD$;$K=F;J=$9<==G=<$DH;>$C@JBD;<:$F;J=C$B>

K<;LF=>D=G$H;9BD;D$72BLE<=$]\UV87!;9J=$]\]5$J@I;DB@>$=KK=ID8?$$1@JBD;<:$F;J=C$S=<=$F@<=

CEII=CCKEJ$B>$<=I<EBDB>L$K=F;J=C$SH=>$DH=$/=BLH9@<$F@M=F=>D$<EJ=$S;C$B>$=KK=ID$DH;>

SH=>$DH=$<EJ=$S;C$*;>G@F$7!;9J=$]\]5$F@M=$=KK=ID85$;>G$SBDH@ED$F;D<BZ$=KK=IDC$DH;>$SBDH

F;D<BZ$=KK=IDC$7!;9J=$]\]5$F;D<BZ$=KK=ID8$72BLE<=$]\UV8?$$3>G=<$DH=$*;>G@F$<EJ=5$9=DS==>

]^\gW_$@K$C@JBD;<:$F;J=C$B>$K<;LF=>D=G$H;9BD;D$GB=G$SBDH@ED$<=I<EBDB>L$;$K=F;J=$72BLE<=

]\UV8?$$3>G=<$DH=$/=BLH9@<$<EJ=5$GBKK=<=>I=C$9=DS==>$K<;LF=>D=G$;>G$I@>DBLE@EC

A@AEJ;DB@>C$S=<=$J=CC$=ZD<=F=$7!;9J=$]\]5$J@I;DB@>{F@M=$=KK=ID8?$$*E>C$ECB>L$DH=

*;>G@F$F@M=F=>D$<EJ=$;>G$DH=$F@CD$C=M=<=$F;D<BZ$=KK=ID$71D=AVW8$I<=;D=G$DH=$J;<L=CD

GBKK=<=>I=C$9=DS==>$A@AEJ;DB@>C$B>$K<;LF=>D=G$;>G$I@>DBLE@EC$H;9BD;D5$;>G$F@CD$IJ@C=J:

F;DIH=G$KB=JG$@9C=<M;DB@>C?

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0(1,311()/

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GBCA=<C;J$F@M=F=>D$<EJ=$;>G$D@$DH=$GBCA=<C;J$F@<D;JBD:$<EJ=?$$2E<DH=<F@<=5$;JJ$GBKK=<=>I=C

B>$A@AEJ;DB@>$CB[=$9=DS==>$K<;LF=>D=G$;>G$I@>DBLE@EC$H;9BD;D$S=<=$;$GB<=ID$<=CEJD$@K

GBKK=<=>I=C$B>$K=F;J=$GBCA=<C;J$CEII=CC$;>G$<=I<EBDF=>D5$<;DH=<$DH;>$9B<DH$;>G$G=;DH$<;D=C?

2<@F$DHBC$($G<;S$C=M=<;J$I@>IJECB@>C$SHBIH$($SBJJ$GBCIECC$9=J@S?$$2B<CD5$DH=$*;>G@F$<EJ=

9=DD=<$F;DIH=G$KB=JG$@9C=<M;DB@>C5$CELL=CDB>L$DH;D$DH=$GBCD<B9EDB@>$@K$S@@GJ;>G$H;9BD;D

I;>$C=M=<=J:$<=CD<BID$DH=$C=;<IH$=KKBIB=>I:$@K$%<@S>$!<==I<==A=<C?$$1=I@>G5$DH=$C=M=<=

F;D<BZ$=KK=ID$F@CD$IJ@C=J:$F;DIH=G$KB=JG$@9C=<M;DB@>C5$CELL=CDB>L$DH;D$%<@S>

!<==I<==A=<C$F;:$=ZA=<B=>I=$HBLH$F@<D;JBD:$B>$>@>\S@@GJ;>G$H;9BD;D?$$!HB<G5$DH=$<=J;DBM=

B>KJE=>I=$@K$GBCA=<C;J$@>$A@AEJ;DB@>$CB[=$S;C$G=A=>G=>D$@>$D=<<BD@<:$CA;DB;J$;<<;>L=F=>D?

2B>;JJ:5$F@<=$<=C=;<IH$BC$>==G=G$I@>I=<>B>L$F@M=F=>D$A;DD=<>C$;>G$GBCA=<C;J$G=IBCB@>C

B>$@<G=<$K@<$F@G=J$<=CEJDC$D@$9=$F=;>B>LKEJJ:$;AAJB=G$D@$I@>C=<M;DB@>$;IDB@>C$CEIH$;C

<=C=<M=$G=CBL>$;D$;$J;>GCI;A=$CI;J=?

Movement Rules - !H=$BFA@<D;>I=$@K$D=<<BD@<:$IJEFAB>L$B>$DH=$F@G=J$BC$EJDBF;D=J:

JB>P=G$D@$C=;<IH$CD<;D=LB=C?$$,JEFAB>L$@K$D=<<BD@<B=C$;JJ@SC$GBCA=<C;J$CEII=CC$;>G

<=I<EBDF=>D$E>G=<$CBFAJ=$C=;<IH$<EJ=C5$SH=<=;C$GBCA=<C;J$CEII=CC$;F@>L$>@>\IJEFA=G

D=<<BD@<B=C$<=NEB<=C$I@FAJ=Z$C=;<IH$<EJ=C5$CEIH$;C$/=BLH9@<?$$!E<>=<$=D$;J?$7UXXY8$<=;IH=G

CBFBJ;<$I@>IJECB@>C$B>$;>$;>;J@L@EC$CBFEJ;DB@>$@K$K@<;LB>L$@K$E>LEJ;D=C$B>$.=JJ@SCD@>=

/;DB@>;J$4;<P?$$!H=:$K@E>G$DH;D$DH=$;<<;>L=F=>D$@K$<=C@E<I=C$7IJEFA=G$@<$<;>G@F8

F;DD=<=G$@>J:$SH=>$F@M=F=>D$S;C$JBFBD=G?$$!HBC$F;DIH=C$B>DEBDBM=$=ZA=ID;DB@>C$DH;D

C=;<IHB>L$K@<$;$<=C@E<I=$7CEIH$;C$K@@G$@<$;$D=<<BD@<:8$BC$=;CB=<$SH=>$DH;D$<=C@E<I=$BC

IJEFA=G?$$.=D5$BK$;>$@<L;>BCF$I;>$=FAJ@:$;>$=KKBIB=>D$@<$I@FAJ=Z$C=;<IH$CD<;D=L:5$DH=

GBCD<B9EDB@>$@K$<=C@E<I=C$9=I@F=C$J=CC$BFA@<D;>D?

Mortality Rules$\$(>$DHBC$F@G=J5$F;D<BZ$I@FA@CBDB@>$;KK=ID=G$GBCA=<C;J$GBCD;>I=C

;>G$CEII=CC5$J=;GB>L$D@$BFA;IDC$@>$A@AEJ;DB@>$A=<CBCD=>I=?$$&JDH@ELH$J@>L\GBCD;>I=

GBCA=<C;J$=M=>DC$@IIE<<=G$;D$K;B<J:$J@S$K<=NE=>IB=C$=M=>$;F@>L$I@>DBLE@EC$A@AEJ;DB@>C5

F;D<BZ$=KK=IDC$KE<DH=<$G=I<=;C=G$DH=$K<=NE=>I:$@K$J@>L\GBCD;>I=$F@M=F=>DC$;>G$DHEC

G=I<=;C=G$K=F;J=$<=I<EBDF=>D$;>G$A@AEJ;DB@>$CB[=?$$($@>J:$F@G=J=G$DS@$F;D<BZ$=KK=IDC$;>G

cU

F@<=$=ZD<=F=$=KK=IDC$;<=$A@CCB9J=?$$2@<$=Z;FAJ=5$IJ=;<=G$J;>G$F;:$9=$;$I@FAJ=D=$9;<<B=<

D@$%<@S>$!<==I<==A=<$F@M=F=>DC5$<;DH=<$DH;>$;$L<=;D$<BCP?$$&C$;$9;<<B=<5$IJ=;<=G$J;>G$F;:

;KK=ID$=M=>$CH@<D\GBCD;>I=$F@M=F=>DC?

!H=<=$BC$;>=IG@D;J$=MBG=>I=$DH;D$>@>\K@<=CD=G$H;9BD;D$BC$;$9;<<B=<$D@$F@M=F=>D?$(>

UXXg5$;$KB=JG$;CCBCD;>D$;>G$($KBDD=G$V$H=JA=<$F;J=C$SBDH$<;GB@\D<;>CFBDD=<C$;>G$F@M=G$DH=F

K<@F$DH=B<$<=CA=IDBM=$<=CBG=>D$D=<<BD@<:$D@$D=<<BD@<B=C$V$PF$;S;:?$$)>=$F;J=$S;C$F@M=G

CEIH$DH;D$DH=$CD<;BLHD\JB>=$GBCD;>I=$9=DS==>$DH=$DS@$D=<<BD@<B=C$S;C$I@FA@C=G$@K

S@@GJ;>GC?$$!H=$@DH=<$F;J=$S;C$F@M=G$CEIH$DH;D$DH=$CD<;BLHD\JB>=$GBCD;>I=$9=DS==>$DH=

DS@$D=<<BD@<B=C$S;C$I@FA@C=G$@K$CA;<C=$D<==C$;>G$IJ=;<=G$J;>G5$:=D$;$IB<IEBD@EC$<=DE<>$A;DH

DH@ELH$S@@GJ;>GC$S;C$A@CCB9J=?$$($;>DBIBA;D=G$DH;D$=;IH$9B<G$S@EJG$BFF=GB;D=J:$<=DE<>$D@

BDC$D=<<BD@<:$SBDHB>$H@E<C?$$0EL$K@<;:C5$S=$@9C=<M=G$%<@S>$!<==I<==A=<C$F@MB>L$;D

J=;CD$^WWF$B>$J=CC$DH;>$H;JK$;>$H@E<?$$.=D5$>=BDH=<$9B<G$S;C$;9J=$D@$<=DE<>$H@F=$<=;GBJ:?$$(>

DH=$KB<CD$I;C=5$DH=$F;J=$J@CD$HBC$<;GB@\D<;>CFBDD=<$DH=$G;:$;KD=<$<=J=;C=$GEL$;$KBLHD$SHBJ=

B>D<EGB>L$@>$;>@DH=<$%<@S>$!<==I<==A=<$D=<<BD@<:?$$6=$G=D=ID=G$DH=$9B<G$9;IP$@>$HBC

D=<<BD@<:$]$G;:C$;KD=<$<=J=;C=?$$"=$S;C$<=J=;C=G$B>$;$>;<<@S5$JB>=;<$H;9BD;D$;>G$DH=<=K@<=

H;G$;>$=M=>$IH;>I=$@K$C=J=IDB>L$DH=$I@<<=ID$GB<=IDB@>$;C$;$<=DE<>$A;DH?$$(>$DH=$C=I@>G$I;C=5

DH=$9B<G$K@<;:=G$EA$D@$U$PF$B>$=M=<:$GB<=IDB@>$DH;D$S;C$I@FA<BC=G$@K$S@@GJ;>GC5$9ED$H=

>=M=<$=>D=<=G$>@>\S@@GJ;>G$H;9BD;D?$$)>$DH=$K@E<DH$G;:$;KD=<$<=J=;C=5$S=$I@EJG$>@D$J@I;D=

DH=$9B<G$;>G$H=$>=M=<$<=DE<>=G$D@$HBC$@<BLB>;J$D=<<BD@<:?$$!H=C=$@9C=<M;DB@>C$<=M=;J$DH=

JBFBD=G$F@M=F=>D$9=H;MB@<$@K$F;J=$%<@S>$!<==I<==A=<C?$$3>K@<DE>;D=J:5$($I;>$>@D

G=D=<FB>=$SH=DH=<$DHBC$B>K=<=>I=$;AAJB=C$D@$K=F;J=C5$SH@C=$GBCA=<C;J$9=H;MB@<$BC$F@<=

I<BDBI;J$D@$A@AEJ;DB@>$G:>;FBIC$DH;>$DH;D$@K$F;J=C?

Dispersal success and population size - )DH=<$CBFEJ;DB@>C$7=?L?5$QIR=JM=:$=D$;J?

UXXY5$+=DIH=<$=D$;J?$UXXc8$;JC@$K@E>G$DH;D$GBCA=<C=G$D=<<BD@<B=C$<=CEJD=G$B>$J@S=<

A@AEJ;DB@>$L<@SDH$DH;>$IJEFA=G$D=<<BD@<B=C$GE=$D@$F@<=$=KKBIB=>D$C=;<IHB>L$9:$GBCA=<C=<C

;F@>L$IJEFA=G$D=<<BD@<B=C?$$!H=C=$CDEGB=C$BJJECD<;D=G$DH=$BFA@<D;>I=$@K$GBCA=<C;J$9=H;MB@<

D@$A@AEJ;DB@>$G:>;FBIC?$$"@S=M=<5$GBCA=<C;J$9=H;MB@<$BC$>@D$;JS;:C$BFA@<D;>D$D@

A@AEJ;DB@>$A=<CBCD=>I=?$$1@EDH$7UXXX8$CH@S=G$DH;D$=<<@<C$B>$=CDBF;DB>L$GBCA=<C;J$CEII=CC

;<=$>@D$=NEBM;J=>D$D@$=<<@<C$B>$A<=GBID=G$A@AEJ;DB@>$MB;9BJBD:$9=I;EC=$GBCA=<C;J$CEII=CC$GBG

>@D$;JS;:C$H;M=$;>$=KK=ID$@>$A@AEJ;DB@>$A=<CBCD=>I=$@<$A;DIH$@IIEA;>I:?$$4EJJB;F$=D$;J?

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DH;>$@DH=<$G=F@L<;AHBI$K;ID@<C?

(>$%<@S>$!<==I<==A=<$CBFEJ;DB@>C5$A@AEJ;DB@>C$DH;D$B>I<=;C=G$NEBIPJ:$H;G$;

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CJ@SJ:5$E>G=<$;JJ$F@M=F=>D$<EJ=C5$G=CABD=$DH=$K;ID$DH;D$G=IJB>B>L$A@AEJ;DB@>C$D=>G=G$D@

H;M=$J@>L=<$F;ZBFEF$GBCA=<C;J$GBCD;>I=C?$$(D$;AA=;<C$DH;D$CH@<D\GBCD;>I=$F@M=F=>DC$I;>

H;M=$;$L<=;D=<$=KK=ID$@>$A@AEJ;DB@>$9=H;MB@<$DH;>$J@>L\GBCD;>I=$F@M=F=>DC?$$/B>=D:

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DHBC5$BD$BC$CE<A<BCB>L$DH;D$F;D<BZ$=KK=IDC$B>KJE=>I=G$A@AEJ;DB@>$9=H;MB@<$9=I;EC=$DH=$F;D<BZ

BC$F@<=$JBP=J:$D@$;KK=ID$J@>L\GBCD;>I=$F@M=F=>DC$DH;>$CH@<D\GBCD;>I=$F@M=F=>DC?$$$(>

@DH=<$S@<GC5$($F;:$H;M=$E>G=<=CDBF;D=G$DH=$BFA@<D;>I=$@K$J@>L\GBCD;>I=$GBCA=<C;J

9=I;EC=$DH=$HBLH$IJEFAB>L$@K$D=<<BD@<B=C$I<=;D=C$;$A<=G@FB>;>I=$@K$CEII=CCKEJ$CH@<D\

GBCD;>I=$F@M=F=>DC?$$(K$D=<<BD@<B=C$S=<=$J=CC$IJEFA=G5$J@>L=<\GBCD;>I=$F@M=F=>DC$S@EJG

9=$F@<=$I@FF@>5$I<=;DB>L$;$CBDE;DB@>$SH=<=$F;D<BZ$=J=F=>DC$I@EJG$F@<=$CD<@>LJ:$;KK=ID

GBCA=<C;J$CEII=CC?$$3>G=<$;>@DH=<$CI=>;<B@5$BK$HBLH$CD@IH;CDBIBD:$S=<=$D@$I;EC=$J@I;J

A@AEJ;DB@>$=ZDB>IDB@>C5$J@>L\GBCD;>I=$F@M=F=>DC$S@EJG$9=$>=I=CC;<:$K@<$<=LB@>;J

A@AEJ;DB@>$CD;9BJBD:?

Search rules - Q:$KB>GB>LC$HBLHJBLHD$DH=$>==G$K@<$F@<=$<=C=;<IH$I@>I=<>B>L

F@M=F=>D$A;DD=<>C$;>G$GBCA=<C;J$G=IBCB@>C$B>$@<G=<$K@<$F@G=J$KB>GB>LC$D@$9=$F=;>B>LKEJJ:

;AAJB=G$D@$I@>C=<M;DB@>?$$Q@CD$<=C=;<IH$I@>I=<>B>L$C=;<IH$A;DHC$H;C$9==>$@>$B>M=<D=9<;D=C

7C==$6B=>C$;>G$QBJ>=$UXcX5$!E<IHB>$UXXc85$9ED$BD$BC$E>IJ=;<$H@S$<=J=M;>D$@<$CBFBJ;M=<D=9<;D=$F@M=F=>D$<EJ=C$;<=$D@$M=<D=9<;D=C5$A;<DBIEJ;<J:$9B<GC?$$%B<GC$;AA=;<$HBLHJ:

F@9BJ=$E>G=<$C@F=$IB<IEFCD;>I=C5$CEIH$;C$GEL$FBL<;DB@>5$9ED$=M=>$FBL<;>DC$;AA=;<

I@>CD<;B>=G$9:$C@F=$J;>GCI;A=$K=;DE<=C$GEL$CH@<D\GBCD;>I=$F@M=F=>DC$70=C<@IH=<C

;>G$";>>@>$UXXh5$1D,J;B<$=D$;J?$UXXc8?$$6BDH@ED$F@<=$B>K@<F;DB@>$@>$H@S$9B<GC$F@M=

DH<@ELH$I@FAJ=Z$J;>GCI;A=C5$DH=$G=L<==$D@$SHBIH$F;D<BZ$I@FA@CBDB@>$I@>D<B9ED=C$D@

BC@J;DB@>$=KK=IDC$SBJJ$<=F;B>$E>I=<D;B>?

Territory arrangement - !HBC$F@G=J5$JBP=$F@CD$@DH=<C5$I<=;D=G$BC@J;DB@>$=KK=IDC

DH<@ELH$;$I@F9B>;DB@>$@K$F@M=F=>D$<EJ=C$;>G$D=<<BD@<:$CA;IB>L?$$!H=<=K@<=5$DH=$<=J=M;>I=

@K$DH=$<=CEJDC$SBJJ$G=A=>G$>@D$@>J:$@>$@E<$P>@SJ=GL=$@K$C=;<IH$A;DD=<>C5$9ED$;JC@$@>$@E@SJ=GL=$@K$D=<<BD@<:$GBCD<B9EDB@>5$SHBIH$H;C$<=I=BM=G$JBDDJ=$;DD=>DB@>?$$(>$DHBC$F@G=J5

cY

D=<<BD@<:$CA;DB;J$;<<;>L=F=>D$S;C$>@D$9;C=G$@>$DH=$;IDE;J$I@@<GB>;D=C$@K$F@>BD@<=G

D=<<BD@<B=C5$<;DH=<$D=<<BD@<:$J@I;DB@>C$S=<=$G=D=<FB>=G$9:$H;9BD;D$K<;LF=>D;DB@>$A;DD=<>C$;C

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Initial Conditions_D=<<BD@<:$I@M=<;L= /& UW_

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J=DD=<C$B>GBI;D=$CBL>BKBI;>D$<=CEJDC?

Currency Effect DF F Value Pr > F

Q=;>$A@AEJ;DB@>$CB[=$;D$.]W

Location $U$$$$$VgV?]] x?WWWU

Move U$$$$$V]g?g] x?WWWU

Matrix $$ V$$$$$Yc?hg x?WWWU

Location*Move$$ U$$$$$$YW?XX x?WWWU

Location*Matrix$$ V$$$$$$^?Xh$ W?WWYW

Move*Matrix$$$$ V$$$$$$UU?^V$ x?WWWU

+@I;DB@>{Q@M={Q;D<BZ V$$$$$$$$W?WW$ U?WWWW

Q=;>$F;ZBFEF$GBCD;>I=

+@I;DB@>$$ U$$$$$$$$W?]^ W?^WWc

Move $$$$ U$$$$$$$UU?cc W?WWWh

Matrix $ V$$$$$]^^?gX x?WWWU

+@I;DB@>{Q@M=$ U$$$$$$$$U?^V W?VUc]

Location*Matrix V$$$$$$$UV?^] x?WWWU

Q@M={Q;D<BZ$ V$$$$$$$$W?gc W?^Wg^

+@I;DB@>{Q@M={Q;D<BZ V$$$$$$$$$W?WU W?XcgV

X^

!;9J=$]\]$I@>DB>E=G?

Currency Effect DF F Value Pr > F

_$J@>L\GBCD;>I=$F@M=C

Location U$$$$$$Vc]?]Y x?WWWU

Move $$ U$$$$$$$UYV?cW x?WWWU

Matrix $ V$$$$$UWVW?UV x?WWWU

Location*Move$ U$$$$$$$g]?WU x?WWWU

Location*Matrix$$ V$$$$$$$gW?Wg x?WWWU

Move*Matrix$$ V$$$$$$$VV?UW x?WWWU

Location*Move*Matrix V$$$$$$$$]?cW W?WWXU

_$F;J=C$DH;D$<=I<EBD$;$K=F;J=

Location$ U$$$$$$YXV?]c x?WWWU

Move$ U$$$$$Vcg?]U$ x?WWWU

Matrix $$ V$$$$$^Y?U] x?WWWU

Location*Move$$ U$$$$VU?YW x?WWWU

Location*Matrix $ V$$$$$$$Y?XV W?WVUV

Q@M={Q;D<BZ$$$$ V$$$$$$$V?VW W?UUY]

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UWW

020

4060

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om

eter

sN

UWU

Q;J=

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4$y$?U^

%

UW]

0

1

2

3

4

5

6

7

8

0 50 100 150 200 250 300

mor

talit

y m

ultip

lier

A

0

5

10

15

20

25

0 50 100 150 200 250 300

relative habitat quality

mor

talit

y m

ultip

lier

B

UW^

-0.2

5-0

.2-0

.15

-0.1

-0.0

50

0.05

Pro

porti

onal

Sen

sitiv

ity o

f Lam

bda

Mal

e D

ispe

rsal

Pro

b

Fbr

eede

r Mor

talit

y

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sper

ser M

orta

lity

Ffle

dgin

g M

orta

lity

Mbr

eede

r Mor

talit

y

Mso

litar

y M

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lity

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Mor

talit

y

Mdi

sper

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lity

Mfle

dglin

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lity

UWg

Random, Constant

0

50

100

150

200

250

300

0 10 20 30 40

# pa

irs

Random, Cubic7

0

50

100

150

200

250

300

0 10 20 30 40

# pa

irs

Random, Step20

0

50

100

150

200

250

300

0 10 20 30 40

Year

# pa

irs

UWh

Neighbor, Constant

0

50

100

150

200

250

300

0 10 20 30 40

# pa

irs

Neighbor, Cubic7

0

50

100

150

200

250

300

0 10 20 30 40

# pa

irs

Neighbor, Step20

0

50

100

150

200

250

300

0 10 20 30 40

Year

# pa

irs

UWc

050100

150

200

250

300

350

N-C

onst

ant

N -

C7

N -

S20

R-C

onst

ant

R -

C7

R -

S20

mean # pairs at Y40C

ontig

uous

Fra

gmen

ted

UWX

WV]gcUWUVU]UgUc

/$\$,@>CD;>D

/$\$,E9BIh

/$\$1D=AVW

*$\$,@>CD;>D

*$\$,E9BIh

*$\$1D=AVW

% cross-overs

UUW

051015202530354045

N -

Con

stan

tN

- C

ubic

7N

- S

tep2

0R

- C

onst

ant

R -

Cub

ic7

R -

Ste

p20

maximum distance (km)

UUU

05101520253035

N -

Con

stan

tN

- C

ubic

7N

- S

tep2

0R

- C

onst

ant

R -

Cub

ic7

R -

Ste

p20

% long moves

UUV

2030405060708090100

N -

Con

stan

tN

- C

ubic

7N

- S

tep2

0R

- C

onst

ant

R -

Cub

ic7

R -

Ste

p20

% solitaries recruit female

Con

tiguo

us

Fra

gmen

ted

UUY

CHAPTER 5. HABITAT AND LANDSCAPE EFFECTS ON BROWNTREECREEPER DISTRIBUTION: INDEPENDENT EFFECTS OFWOODLAND LOSS AND FRAGMENTATION AT MULTIPLESCALES

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4060

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11.5

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0 5 10 15 20 25 30 35 40

%woods (4.5km scale)

2nd

PC

-2

-1

0

1

2

3

4

0 10 20 30 40 50 60

%woods (2.0 km scale)

2nd

PC

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0

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0.6

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1

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0 20 40 60 80 100

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GR

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0.6

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1

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0 20 40 60 80 100

#cavities per hectare

GR

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0

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U]V

CHAPTER 6. HABITAT VARIATION AND COOPERATIVEBREEDING OF THE AUSTRALIAN BROWN TREECREEPER

&%1!*&,!

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Statistical Analyses8?

Statistical Analyses$\$($EC=G$JB>=;<$<=L<=CCB@>$D@$=Z;FB>=$DH=$L=>=<;J$<=J;DB@>CHBA

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*#13+!1

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0(1,311()/

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H=ID;<=$D<==$K;ID@<C?$$&JJ$K;ID@<C$S=<=$L=>=<;D=G$9:$AIBA;J$I@FA@>=>DC$;>;J:CBC$7/$y

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';<B;9J=$C=D$U

4,\& 4,\% 4,\, 4,\0

#BL=>M;JE= V?U U?^ U?Y U?U

4<@A@<DB@>$@K$ W?YW W?VV W?UX W?U^

Variable

%;<=$L<@E>G \W?Wh W?Vh -0.77 W?V]

+@S$L<;CC -0.65 \W?WWV W?U] \W?VV

!;JJ$L<;CC 0.55 \W?YY W?VY W?YY

*@IPC W?V^ 0.45 W?V^ -0.63

+@LC \W?U] W?Y] 0.46 0.44

0S;<K$CH<E9 \W?V^ W?YY W?VU 0.44

!;JJ$CH<E9 W?Y^ 0.63 \W?WX W?W^

';<B;9J=$C=D$V

4,U 4,V 4,Y#BL=>M;JE= U?^ U?Y U?W

4<@A@<DB@> W?YW W?Vg W?VW

Variable

+BM=$D<==C \W?h 0.66 0.56

%@Z=C -0.49 0.54 \W?Vh

-EFC 0.60 W?YY W?Vg

1>;LC 0.56 \W?U^ \W?Wh

1DL:9;<PC \W?VX -0.42 0.74

UgU

!;9J=$g\]?$$+B>=;<$F@G=JC$I<=;D=G$9:$CD=ASBC=$FEJDBAJ=$<=L<=CCB@>C$@K$9B<G$M;<B;9J=C

7!;9J=$g\V8$@>$M;<B;9J=$C=D$U$;>G$V$7!;9J=$g\Y8?$$+B>=;<$F@G=JC$I<=;D=G$9:$CD=ASBC=

FEJDBAJ=$<=L<=CCB@>C$@K$0#/1(!.$;>G$%*))0$@>$M;<B;9J=$C=D$U$;>G$V$7!;9J=$g\Y8?$$/@

H;9BD;D$M;<B;9J=C$=ZAJ;B>=G$M;<B;DB@>$B>$L<@EA$CB[=$@<$2+#0-#?

*=CA@>C= *=L<=CC@<$ #CDBF;D=$7}1#8 2 ;Gi*V 4$w$!

';<B;9J=$1=D$U$e$V$7A=<$H=ID;<=$=CDBF;D=C8

0#/1(!.$ ,&' W?WV$7W?WU8 h?Xg W?VV W?WU

%*))0$ ,&' W?WV$7W?WU8 c?Xh W?]Y W?WWY

4,V \W?Yh$7W?UU8 UU?^Y

UgV

2(-3*#$,&4!()/1

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D=<<BD@<B=C5$7&8$F=;CE<=G$;C$DH=$>EF9=<$@K$KJ=GLJB>LC$A<@GEI=G$A=<$:=;<$;>G$7%8$F=;CE<=G

;C$DH=$>EF9=<$@K$KJ=GLJB>LC$K<@F$CEII=CCKEJ$>=CDC$A=<$:=;<?$$%J;IP$IB<IJ=C$B>GBI;D=$DH=

F=;>$;>G$M=<DBI;J$JB>=C$B>GBI;D=$}$U$CD;>G;<G$G=MB;DB@>?

2BLE<=$g\V?$$1BFAJ=$JB>=;<$<=L<=CCB@>$@K$L<@EA$CB[=$@>$<=A<@GEIDBM=$CEII=CC$;KD=<

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DH=$>EF9=<$@K$KJ=GLJB>LC$A<@GEI=G?

2BLE<=$g\Y?$$1BFAJ=$JB>=;<$<=L<=CCB@>$@K$0#/1(!.$7L<@EA$CB[=$B>G=A=>G=>D$@K$D=<<BD@<:

CB[=8$@>$I;MBD:$G=>CBD:$7/$y$V^8?

2BLE<=$g\]?$$1I;DD=<$AJ@DC$@K$%*))0$;L;B>CD$I;MBDB=C$A=<$H=ID;<=$7/$y$V^8?

2BLE<=$g\^?$$1I;DD=<$AJ@D$@K$%*))0$;L;B>CD$9;C;J$;<=;$@K$D<==C$A=<$H=ID;<=$7/$y$V^8?

2BLE<=$g\g?$$!H=$H:A@DH=DBI;J$GBCD<B9EDB@>$@K$I;MBD:$G=>CBD:$;>G$KBD>=CC$E>G=<$7&8$DH=

I<BDBI;J\<=C@E<I=$F@G=J5$7%8$DH=$M;<B;>I=$F@G=J5$;>G$7,8$DH=$F;<LB>;J\H;9BD;D$F@G=J?

UgY

0

0.5

1

1.5

2

2.5

3

3.5

4

1 2 3 4 5

Group size

Bro

od

siz

e

Brood size = 1.29 + 0.23GS adjR2 = .04 P = 0.05

A

-0.5

0

0.5

1

1.5

2

2.5

3

3.5

4

1 2 3 4 5 6

Group size

# fl

edg

ling

s p

er y

ear

Fledges = 0.14 + 0.44GS adjR2 = 0.10 P = 0.0003B

Ug]

0

0.5

1

1.5

2

2.5

3

3.5

1 1.5 2 2.5 3 3.5 4 4.5 5 5.5

Mean group size

mea

n #

fle

dg

ling

smean fledged = -0.05 + 0.50meanGS adjR2 = 0.24 P = 0.007

Ug^

0

0.5

1

1.5

2

2.5

3

3.5

0 10 20 30 40 50 60 70 80

# cavities per hectare

Den

sity

Density = 0.49 + 0.02cav adjR2 = 0.22 P = 0.01

Ugg

0

0.5

1

1.5

2

2.5

3

3.5

4

4.5

0 10 20 30 40 50 60 70 80

CAV

BR

OO

D

Ugh

0

0.5

1

1.5

2

2.5

3

3.5

4

4.5

0 10 20 30 40 50

TREE

BR

OO

D

Ugc

-1

-0.5

0

0.5

1

1.5

2

2.5

0 50 100 150 200 250 300 350

cavity density

fitn

ess

A

-1

-0.5

0

0.5

1

1.5

2

2.5

3

0 50 100 150 200 250 300 350

cavity density

fitn

ess

B

-1

-0.5

0

0.5

1

1.5

2

2.5

0 50 100 150 200 250 300 350

cavity density

fitn

ess

C

UgX

CHAPTER 7. CONCLUSIONS1./!"#1(1

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Cirriculum Vitae

Caren B. Cooper, [email protected]=AD?$@K$%B@J@L:5$!EKDC$3>BM=<CBD:5$Q=GK@<G5$Q&$WVU^^

Education

UXX^\VWWW 'B<LB>B;$4@J:D=IH>BI$(>CDBDED=$e$1D;D=$3>BM=<CBD:4H?0?$1=AD=F9=<$VWWW0=A;<DF=>D$@K$%B@J@L:5O=KK=<:$6;JD=<C5$,@FFBDD==$,H;B<

UXXU\UXXY$$$$$$$3>BM=<CBD:$@K$6:@FB>LQ?1?$0=I=F9=<$UXXY0=A;<DF=>D$@K$l@@J@L:$;>G$4H:CB@J@L:5$-4&$]?W1D;>J=:$&>G=<C@>5$,@FFBDD==$,H;B<

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UXch\cc "EF9@JGD$1D;D=$3>BM=<CBD:5$,&$7/;DB@>;J$#ZIH;>L=$1DEG=>D8

Independent and Collaborative Research Experience

Post Doctoral Research Associate5$0=A;<DF=>D$@K$%B@J@L:5$!EKDC$3>BM=<CBD:5$Q& 2000

!H=$B>KJE=>I=$@K$DBF9=<$H;<M=CD$A;DD=<>C$@>$9B@GBM=<CBD:$ECB>L$;$CBFEJ;DB@>$F@G=J?

Graduate Research Associate, 0=A;<DF=>D$@K$%B@J@L:5$'B<LB>B;$4@J:D=IH>BI 1995-2000

(>CDBDED=$e$1D;D=$3>BM=<CBD:5$'&0=CBL>=G$;>G$I;<<B=G$@ED$<=C=;<IH$@>$%<@S>$!<==<==A=<C?$$2B=JG$S@<PB>IJEG=G$9;>GB>L$;GEJDC$;KD=<$I;ADE<=$B>$FBCD$>=CDC5$9;>GB>L$>=CDJB>LC5@9C=<MB>L$L<@EA$B>D=<;IDB@>C5$F@>BD@L$>=CD$CEII=CC5$F=;CELH;9BD;D$M;<B;9J=C5$;>G$CEA=<MBCB>L$KB=JG$;CCBCD;>DC?$$3C=G$Q1#ZI=J51&15$;>G$&<I'B=S$D@$@<L;>B[=$;>G$;>;J:[=$G;D;?$$3C=G$&<I(>K@$D@GBLBDB[=$J;>GCI;A=$I@M=<D:A=C?$$3C=G$'BCE;J$%;CBI$D@$<E>$CBFEJ;DB@>F@G=J$@K$%<@S>$!<==I<==A=<$A@AEJ;DB@>$G:>;FBIC?

Research Associate5$3>BM=<CBD:$@K$(G;H@5$(0 $$$$$$$$$$$$$$$$1994

,@>GEID=G$ABJ@D$CDEG:$B>M=CDBL;DB>L$K@<;LB>L$J@I;DB@>C$@K$!E>G<;1S;>C$;D$;$FBL<;DB@>$CD@A\@M=<$CBD=$B>$/@<DH=<>$(G;H@$SBDH$J=;GI@>D;FB>;DB@>?

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Graduate Research Associate5$0=A;<DF=>D$@K$l@@J@L:$;>G$4H:CB@J@L:5 1991-1993

3>BM=<CBD:$@K$6:@FB>L5$6.0=CBL>=G$;>G$I;<<B=G$@ED$<=C=;<IH$@>$DH=$EC=$@K$I@>CD<EID=G$S=DJ;>GC9:$G;99JB>L$GEIPC?$$2B=JG$S@<P$B>IJEG=G$C;FAJB>L$CE9F=<L=>DM=L=D;DB@>5$<=I@<GB>L$G=D;BJ=G$K@<;LB>L$@9C=<M;DB@>C5$CE<M=:B>L$K@<GEIPC5$;>G$CEA=<MBCB>L$KB=JG$;CCBCD;>DC?$$3C=G$Q1#ZI=J5$1&15$;>G2@<D<;>$D@$@<L;>B[=$;>G$;>;J:[=$G;D;?

Field Research Experience

Research Technician5$1FBDHC@>B;>$,@>C=<M;DB@>$;>G$*=C=;<IH$,=>D=<5$'& 1991

1=;<IH=G$K@<$R=>DEIP:$S;<9J=<C5$<=;G$I@J@<$9;>GC5$C=;<IH=G$K@<$>=CDC5I;ADE<=G$;GEJDC$B>$FBCD\>=DC5$;>G$9;>G=G$>=CDJB>L?

Research Technician5$1FBDHC@>B;>$,@>C=<M;DB@>$;>G$*=C=;<IH$,=>D=<5$'& UXXW&CCBCD=G$B>$<=C=;<IH$DH;D$;AAJB=G$9=H;MB@<;J$I@>GBDB@>B>L$D<=;DF=>DCA<B@<$D@$<=J=;C=$B>$DH=$SBJG$@K$1B9=<B;>$A@J=I;DC$;C$;$F@G=J$K@<$9J;IP\K@@D=G$K=<<=D$<=B>D<@GEIDB@>?$$0EDB=C$B>IJEG=G$I;L$K@<$I;ADBM=A@J=I;DC$SHBJ=$BFAJ=F=>DB>L$A<=\<=J=;C=$D<;B>B>L$CEIH$;C$IH;CB>LA@J=I;DC$SBDH$;$G@L$;>G$@SJ5$K==GB>L$A@J=I;DC$JBM=$A<;B<B=$G@LC5$;>GG=;G$A<;B<B=$G@LC?

Field Technician5$31$2BCH$;>G$6BJGJBK=$1=<MBI=5$4E=<D@$*BI@ UXXW,@>GEID=G$G;S>$D@$GECP$@9C=<M;DB@>C$@K$>=CDB>L$4E=<D@$*BI;>$4;<<@DC?

Field Technician5$/1,35$0=A;<DF=>D$@K$l@@J@L:5$/, UXcX!<;AA=G5$D;LL=G5$;>G$<;GB@\D<;IP=G$9J;IP$9=;<C$B>$F@E>D;B>@EC$D=<<;B>GEL$CEFF=<?$$+@I;D=G$G=>C$B>$SB>D=<?

Intern5$1DEG=>D$,@>C=<M;DB@>$&CC@IB;DB@>5$4E=<D@$*BI@ UXcX,@>GEID=G$G;S>$D@$GECP$@9C=<M;DB@>C$@K$>=CDB>L$4E=<D@$*BI;>$4;<<@DC?

Field Technician5$31$2@<=CD$1=<MBI=5$RJ;F;DH$/;DB@>;J$2@<=CD5$,& UXcc,@>GEID=G$G;S>$D@$GECP$@9C=<M;DB@>C$@K$K@<;LB>L$1S;B>C@>`C$H;SPC?!<;AA=G$H;SPC$B>$FBCD\>=CDC$K@<$9;>GB>L$;>G$<;GB@\D;LLB>L?

Undergraduate Intern5$/1,35$0=A;<DF=>D$@K$2@<=CD<:5$/, UXch,@>GEID=G$9B<G$I@E>DC$9:$CBLHD$;>G$C@E>G$B>$I:A<=CC\DEA=J@$CS;FA$B>&J;9;F;?$$1;FAJ=G$M=L=D;DB@>$B>$IED$;>G$E>IED$K@<=CD?

Hack Site Attendant5$/,$6BJGJBK=$*=C@E<I=C$,@FFBCCB@>5$/, UXcg2=G$;>G$@9C=<M=G$:@E>L$4=<=L=$2;JI@>C$A<B@<$D@$;>G$;KD=<$DH=B<<=J=;C=?$$4<@D=ID=G$DH=$CBD=$K<@F$B>D<EG=<C?

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Undergraduate Intern5$/,135$0=A;<DF=>D$@K$l@@J@L:5$/, UXc^!<;AA=G5$D;LL=G5$;>G$<;GB@\D<;IP=G$9J;IP$9=;<C$B>$F@E>D;B>@EC$D=<<;B>GEL$CEFF=<?$$+@I;D=G$9=;<$G=>C$B>$SB>D=<?$$1EA=<MBC=G$#;<DHS;DIHM@JE>D==<C?$$1;FAJ=G$M=L=D;DB@>?

Work-Study Intern5$/,135$0=A;<DF=>D$@K$l@@J@L:5$/, UXc]!=>G=G$L<==>H@EC=$L<;CC=C?$1=;<IH=G$I;DD;BJC$K@<$L<;CCH@AA=<$=LLC?Q=;CE<=G5$F;<P=G5$;>G$G=D=<FB>=G$DH=$C=Z$@K$L<;CCH@AA=<C$;KD=<$=;IHF@JD?

Volunteer5$0EP=$4<BF;D=$,=>D=<5$/, UXc]2=G$;>G$@9C=<M=G$DH=$9=H;MB@<$@K$I;ADBM=$J=FE<C?

Laboratory Experience

Research Technician5$1FBDHC@>B;>$,@>C=<M;DB@>$;>G$*=C=;<IH$,=>D=<5$'& UXXU0<=S$9J@@G$C;FAJ=C$K<@F$I;ADBM=$6HBD=\IH==P=G$4B>D;BJ$GEIPC$;>G<=I@<G=G$AJEF;L=$IH;<;ID=<BCDBIC?$$&CCBCD=G$SBDH$I@JEF>IH<@F;D@L<;AH:$D@$C=A;<;D=$CD=<@BG$H@<F@>=C$7D=CD@CD=<@>=5GBH:G<@D=CD@CD=<@>=5$JED=B>B[B>L$H@<F@>=5$;>G$I@<DBI@CD=<@>=8?&CCBCD=G$B>$DH=$F=;CE<=F=>D$@K$H@<F@>=$I@>I=>D<;DB@>C$9:<;GB@BFFE>@;CC;:?

Teaching Experience

Teaching Assistant5$'B<LB>B;$!=IH5$'& UXXc!;ELHD$V$J;9@<;D@<:$C=IDB@>C$@K$@<>BDH@J@L:?$$!@@P$CDEG=>DC$@>$KB=JGD<BAC$;>G$<=MB=S=G$FEC=EF$CA=IBF=>C?$$4<=A;<=G$;>G$L<;G=GJ;9@<;D@<:$A<;IDBI;J$=Z;FC?$$-<;G=G$S<BDB>L$;CCBL>F=>DC?

Associate Naturalist5$!H=$,@>C=<M;>I:5$2+ $$$UXXW\UXXU!@@P$]DH$;>G$gDH$L<;G=<$@>$KB=JG$D<BAC$D@$9=;IH$;>G$EAJ;>G$H;9BD;DC$;CS=JJ$;C$D@$;$J;9@<;D@<:$;D$!H=$,@>C=<M;>I:?$$-EBG=G$9B<G$S;DIHB>LI;>@=$D<BAC$K@<$;GEJDC$D@$<@@P=<:$BCJ;>GC$;>G$DH<@ELH@ED$F;>L<@M=CS;FAC?$$-EBG=G$@EDB>LC$B>$9@;D$D<BAC$D@$9;<<B=<$BCJ;>GC?$$!;ELHDA<=CIH@@J$A<@L<;FC$SBDH$B>iE<=G$SBJGJBK=$K<@F$<=H;9BJBD;DB@>$I=>D=<?

Volunteer Naturalist5$6=CD=<>$/@<DH$,;<@JB>;$/;DE<=$,=>D=<5$/, UXcX!;ELHD$Y<G$L<;G=$A<@L<;F$B>$DH=$K;JJ$;9@ED$;AAJ=C?$$4<@MBG=GG=F@>CD<;DB@>C$D@$L=>=<;J$AE9JBI$;9@ED$9==C5$DE<DJ=C5$;>G$;FAHB9B;>C$;D,=>D=<?

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Environmental Policy Experience

Research Fellow5$!BA$@K$DH=$QBDD$6;D=<CH=G$,@E>IBJ5$Q( UXX^0=M=J@A=G$IEFEJ;DBM=$BFA;ID$;CC=CCF=>D$A<@D@I@J$K@<$S=DJ;>G$A=<FBD<=MB=S$9:$DH=$0=A;<DF=>D$@K$/;DE<;J$*=C@E<I=C?$$,@FF=>D=G$@>G<=GL=\;>G\KBJJ$A=<FBD$;AAJBI;DB@>C?

Other Experience

GIS Analyst5$!H=$/;DE<=$,@>C=<M;>I:5$*( VWWW,@FAED=G$J;>GCI;A=$F=D<BIC$CE<<@E>GB>L$9B<G$I@E>D$CD;DB@>C$ECB>L$&<I'B=S

Research Grants and Awards

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Publications

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Publications In Review

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Publications In Preparation

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Reports

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Abstracts

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Documents

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Documents

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